New fertile specimens of Archaeopteris roemeriana from Upper (but not uppermost) Famennian strata of Belgium have been studied. Crowded sporangia are borne on non-laminate, three times bifurcate fertile leaves helically arranged around the fertile ultimate axes. Vegetative leaves occur both proximal and distal to the fertile leaves. Heterospory is demonstrated with miospores belonging to the Geminospora-Aneurospora complex and macrospores most probably to Contagisporites genus. The established morphological traits allow us to substantiate a synonymy between A. roemeriana and A. halliana. The apparent dominance of A. halliana on the eastern side of the Acadian mountains during Late Famennian times is discussed.

中文翻译：

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来自比利时上法门阶的始祖鸟：异种孢子、命名法和古生物地理学

已经研究了来自比利时上（但不是最上）法门地层的 Archaeopteris roemeriana 的新肥沃标本。拥挤的孢子囊生长在非层状的三分叉可育叶上，螺旋状排列在可育的最终轴周围。营养叶出现在可育叶的近端和远端。异孢子被证明为小孢子属于双孢子-动脉孢子复合体，而大孢子最有可能属于传染性孢子属。既定的形态特征使我们能够证实 A. roemeriana 和 A.halliana 之间的同义词。讨论了法门晚期 A.halliana 在阿卡迪亚山脉东侧的明显优势。