Several taxa that share similar ascomatal and ascospore characters occur in monotypic or small genera throughout the Sordariomycetidae with uncertain relationships based on their morphology. Taxa in the genera Duradens, Leptosporella, Linocarpon, and Rimaconus share similar morphologies of conical ascomata, carbonised peridia and elongate ascospores, while taxa in the genera Caudatispora, Erythromada and Lasiosphaeriella possess clusters of superficial, obovoid ascomata with variable ascospores. Phylogenetic analyses of 28S large-subunit nrDNA sequences were used to test the monophyly of these genera and provide estimates of their relationships within the Sordariomycetidae. Rimaconus coronatus is described as a new species from New Zealand; it clusters with the type species, R. jamaicensis. Leptosporella gregaria is illustrated and a description is provided for this previously published taxon that is the type species and only sequenced representative of the genus. Both of these genera occur in separate, well-supported clades among taxa that form unsupported groups near the Chaetosphaeriales and Helminthosphaeriaceae. Lasiosphaeriella and Linocarpon appear to be polyphyletic with species occurring in several clades throughout the subclass. Caudatispora and Erythromada represented by single specimens and two putative Duradens spp. have unclear affinities in the Sordariomycetidae.

Taxonomic novelties:Rimaconus coronatus Huhndorf & A.N. Mill., sp. nov.

几种共享相似子囊孢子和子囊孢子特征的分类群出现在整个Sordariomycetidae 的单型或小属中，基于它们的形态具有不确定的关系。Duradens、Leptosporella、Linocarpon和Rimaconus属的分类群具有相似的圆锥形子囊、碳化子囊和细长子囊孢子的形态，而Caudatispora、Erythromada和Lasiosphaeriella属中的分类群具有带有可变子囊孢子的浅卵球形子囊簇。28S 大亚基 nrDNA 序列的系统发育分析用于测试这些属的单系性并提供它们在Sordariomycetidae中的关系的估计值。冠毛螟被描述为来自新西兰的新物种；它与模式种R. jamaicensis聚集在一起。对 Leptosporella gregaria进行了说明，并为这个先前公布的分类群提供了描述，该分类群是该属的模式种和唯一的测序代表。这两个属都出现在独立的、受良好支持的进化枝中，这些分类群在毛球菌目和长丝球虫科附近形成不受支持的群体。Lasiosphaeriella和Linocarpon似乎是多系的，在整个亚纲的几个进化枝中出现了物种。Caudatispora和Erythromada由单个标本和两个假定的Duradens 代表属 在Sordariomycetidae 中具有不清楚的亲和力。

分类新奇：Rimaconus coronatus Huhndorf & AN Mill., sp. 十一月