The phylogenetic relationships amongst the Arminidae were analysed based upon morphological characters of 58 presently described species or nudibranchs, including 35 previously described Arminidae and 20 new species of Dermatobranchus. From the literature review and anatomical examinations, 43 characters were considered for 78 taxa. These characters were polarized using Berthella canariensis as the outgroup taxon and the type species of several other genera identified from recent publications. The resulting phylogeny supports the monophyly of Arminidae, Dermatobranchus, Doridina, and Proctonotidae. The paraphyly of the Arminina is further demonstrated in this study. Two previously described, but poorly known, species of Indo-Pacific Armina are redescribed, Armina magna Baba, 1955 and Armina paucifoliata Baba, 1955. The anatomy and taxonomic status of nine previously described species of Dermatobranchus were examined in this study. The anatomy of Dermatobranchus pustulosus (van Hasselt, 1824) has been overlooked since Bergh (1888) illustrated the radula of van Hasselt's specimen. It is redescribed and its range is extended to several new localities in the western Pacific. Dermatobranchus pulcherrimus Miller & Willan, 1986 is considered here as a new synonym of Dermatobranchus rubidus (Gould, 1852). The following 20 species of Dermatobranchus are new and are described in the present paper: Dermatobranchus albineus sp. nov., Dermatobranchus arminus sp. nov., Dermatobranchus caesitius sp. nov., Dermatobranchus caeruleomaculatus sp. nov., Dermatobranchus cymatilis sp. nov., Dermatobranchus dendonephthyphagus sp. nov., Dermatobranchus diagonalis sp. nov., Dermatobranchus earlei sp. nov., Dermatobranchus fasciatus sp. nov., Dermatobranchus funiculus sp. nov., Dermatobranchus kalyptos sp. nov., Dermatobranchus kokonas sp. nov., Dermatobranchus leoni sp. nov., Dermatobranchus microphallus sp. nov., Dermatobranchus oculus sp. nov., Dermatobranchus phyllodes sp. nov., Dermatobranchus piperoides sp. nov., Dermatobranchus rodmani sp. nov., Dermatobranchus semilunus sp. nov., and Dermatobranchus tuberculatus sp. nov. Eighteen of these new taxa are found in the Indo-Pacific tropics and two are found in temperate South Africa, D. albineus and D. arminus. Unique combinations of morphological characters distinguish these as new species of Dermatobranchus. Several species that are externally similar have radically divergent internal morphology, are members of different clades of Dermatobranchus, and represent cryptic species. Especially important is the radular morphology, which shows remarkable diversity of form, probably related directly to the diversification of feeding of members of this clade on various octocorals.

中文翻译：

---

以前未记录的隐匿物种的多样性和丰度：1814 年对印度-太平洋 Arminidae Rafinesque 的系统发育分析（软体动物：裸鳃类动物），描述了 20 种新的皮鳃类动物

根据目前描述的 58 个物种或裸鳃类动物的形态特征分析了 Arminidae 之间的系统发育关系，其中包括 35 个先前描述的 Arminidae 和 20 个新的皮鳃类动物。从文献综述和解剖学检查来看，78 个分类群考虑了 43 个字符。使用 Berthella canariensis 作为外群分类群和从最近的出版物中确定的其他几个属的模式物种来极化这些特征。由此产生的系统发育支持 Arminidae、Dermatobranchus、Doridina 和 Proctonotidae 的单系。这项研究进一步证明了 Arminina 的附属物。重新描述了两种先前描述但鲜为人知的印度-太平洋 Armina 物种，Armina magna Baba，1955 和 Armina paucifoliata Baba，1955。在这项研究中检查了九种先前描述的皮鳃类动物的解剖结构和分类状态。自从Bergh (1888) 说明了van Hasselt 标本的齿状叶以来，脓疱性皮鳃(van Hasselt, 1824) 的解剖结构就被忽视了。它被重新描述，其范围扩展到西太平洋的几个新地点。Dermatobranchus pulcherrimus Miller & Willan, 1986 在此被认为是 Dermatobranchus rubidus (Gould, 1852) 的新同义词。以下 20 种皮枝属是新的，在本论文中有描述：Dermatobranchus albineus sp。十一月，Dermatobranchus arminus sp。十一月，Dermatobranchus caesitius sp。十一月，Dermatobranchus caeruleomaculatus sp. 十一月，Dermatobranchus cymatilis sp。11 月，Dermatobranchus dendonephthyphagus sp. 十一月，Dermatobranchus diagonalis sp。十一月，Dermatobranchus Earlei sp. 11 月，筋膜皮纹菌属。十一月，Dermatobranchus funiculus sp。11 月，Dermatobranchus kalyptos sp. 11 月，Kermatobranchus kokonas sp. 11 月，莱昂尼皮纹虫 sp. 11 月，Dermatobranchus microphallus sp。十一月，Dermatobranchus oculus sp。11 月，Dermatobranchus phyllodes sp. 十一月，Dermatobranchus piperoides sp。11 月，Dermatobranchus rodmani sp. 11 月，半月皮臂菌属。11 月，和结节皮支菌属。十一月 这些新分类群中有 18 个在印度-太平洋热带地区发现，两个在温带南非发现，D. albineus 和 D. arminus。形态特征的独特组合将它们区分为皮鳃科的新物种。一些外部相似的物种具有完全不同的内部形态，属于不同的皮鳃亚科的成员，并代表神秘物种。特别重要的是放射状形态，它显示出显着的形式多样性，可能与该进化枝成员在各种八珊瑚上的摄食多样化直接相关。