* Corresponding authors

^a Hunan Provincial Engineering and Technology Research Center for Agricultural Microbiology Application, Hunan Institute of Microbiology, Changsha, China
E-mail: liuqingshu@hnwyy.cn

^b Key Laboratory of Veterinary Chemical Drugs and Pharmaceutics, Ministry of Agriculture and Rural, Affairs, Shanghai Veterinary Research Institute, Chinese Academy of Agricultural Sciences, Shanghai 200241, China

^c Helmholtz International Lab for Anti-Infectives, Shandong University-Helmholtz Institute of Biotechnology, State Key Laboratory of Microbial Technology, Shandong University, Qingdao, China
E-mail: bianxiaoying@sdu.edu.cn

^d College of Plant Protection, Hunan Agricultural University, Changsha, China
E-mail: wuchen77@hunau.edu.cn